Beginning with a recap of the coming of the Transformers to Earth and the story of Devastator, the OVA then gets its original story underway, as the Autobots are shown to be in the midst of constructing the powerful "Scramble City," overseen by their newest arrival, Ultra Magnus. When the Decepticons learn of this, their combiner robots are deployed to attack, and a battle between them and their Autobot counterparts ensues, focusing on their "Scramble Power" - the interchangeability of the individual limbs - to the extent that at one point, Breakdown of the Stunticons connects to Superion to damage him. At the OVA's conclusion, Scramble City is activated and assumes its robot mode of Metroplex to rout the Decepticons. However, from the ocean depths, the Decepticons' own city, Trypticon, rises.